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What is a rune?

What is a rune? Here are some definitions.

Noun
  1. A letter, or character, belonging to the written language of various ancient Germanic peoples, especially the Scandinavians and the Anglo-Saxons.
  2. A Finnish or Scandinavian epic poem, or a division of one, especially a division of the Kalevala.
  3. A letter or mark used as mystical or magic symbol.
  4. A verse or song, especially one with mystical or mysterious overtones; a spell or an incantation.
  5. (obsolete) A roun (secret or mystery).
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Examples
Furthermore, several rune alphabets or futharks have been left behind to puzzle scholars.
She paused, staring at the back, one finger gently tracing the black rune that appeared to have been burned into it.
His brother had spent nearly all day and night trying to carve a simple rune into the face of a small amethyst.
One was using recordings I made of rune meditations and mixing bits from each rune into a word spell.
One may also bury a piece of wood inscribed with a protection rune or script.
If I was as drunk as these bottles say I was, I couldn't have cast a rune, let alone a spell.

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