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What is a thresher?

What is a thresher? Here are some definitions.

Noun
  1. (agriculture) Anything or anyone that threshes.
  2. (agriculture, obsolete) A now-obsolete hand tool for threshing, also called a flail.
  3. (agriculture) A modern farm machine for threshing grain, now a part of combine harvesters rather than a separate implement.
  4. (zoology) Any of several large pelagic sharks of the genus Alopias, which have a very long tail; more commonly called thresher sharks. [from 1880s]
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Examples
At that time the thresher had its own small engine and was not pulled by a tractor.
At Weydon fair, this corn thresher got drunk and spoke in an ill-tempered way of his wife.
Peering through my viewfinder, I could scarcely believe what I was seeing, because the pelagic thresher shark is extremely rare.
Once grouped, thresher sharks are thought to use their mighty tails to shock the fish until they are confused, at which point they're eaten.
One enduring memory of this wreck was that of a 4m thresher shark lazily cruising the 10m contour above us as we ascended.
He is one of 90 California fishermen who must use pingers when deploying drift nets for swordfish and thresher shark.

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