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What is a protonema?

What is a protonema? Here are some definitions.

Noun
  1. (botany) A filament of cells that forms following the germination of the spores of mosses and liverworts
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Ultimately, the gametophyte grows from a small bud produced by a cell of the protonema that divides and differentiates.
Erect branches may arise from the protonema, and gemmae may be developed on it.
The spore on germination forms a short filament which soon broadens out into the thalloid protonema.
In Ephemerum also we may probably regard the relation of the small plants to the protonema as a primitive one.
The rhizoids are structurally similar to cells of the protonema, but they lack chlorophyll.
Usually the protonema is filamentous and ceases to be evident after the plants have developed.

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