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What are opercula?

What are opercula? Here are some definitions.

Noun
  1. plural of operculum
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Examples
The opercula of various shells differ in the first place as to their chemical composition.
The third section contains the following genera of land shells with opercula.
The opercula of multivalve shells are composed of two or four pieces, which are called the opercular valves.
The figures represent some of the commoner types of opercula.
Acupuncture at a point commonly used to treat xerostomia is associated with activation of the insula and adjacent opercula.
These fish do not have gills or opercula like most bony fishes.

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