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What is an operculum?

What is an operculum? Here are some definitions.

Noun
  1. A covering flap or lidlike structure in plants and animals, such as a gill cover
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Examples
It is able to withdraw the whole of its body into its shell which is then sealed by an amber-coloured operculum.
The halecostomes, of which Amia is an example, have an opercular dilator muscle originating on the mandible which opens the operculum.
The spine on the operculum is the best way to identify an angelfish and to distinguish it from similar-looking species, such as butterfly fish.
The nape lacks cirri, and the first gill arch attaches to the operculum, the latter characteristic distinguishing Tripterygiidae from Clinidae.
Avicularia are small heterozooids in which the zooecium and operculum form a beak-like, snapping structure that deters small predators.
The chimaeran hyoid is, it turns out, quite happy supporting the operculum and has no interest in the palatoquadrate.

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