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What are milestones?

What are milestones? Here are some definitions.

Noun
  1. plural of milestone
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These are cornerstones, flint stones, millstones, limestone, and milestones.
A metal plate on the milestones says they were cast by Burts the ironmongers in Devizes adding to their local importance.
Furthermore, the Zohar says, Adam is an acronym standing for the three milestones of human history.
Brown's paintings are like milestones marking the distance traveled from the cradle to points far into adulthood.
Elections are not the sole gauge of democracy, but they are, of course, important, substantive milestones.
He wraps the book up tidily with a timeline of the company's development milestones.

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