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What is a grouser?

What is a grouser? Here are some definitions.

Noun
  1. A person who grouses; a grumbler
  2. (nautical) A temporary pile driven into the bottom of a river to hold a boat in position; a spud
  3. (engineering) A protrusion on the surface of a wheel or continuous track segment, intended to increase traction in soil, snow, or other loose material
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Examples
Founders of civilisation, interim riparian cotenants of the River Nile and purveyors of the Cheops Pyramid thrice, I am proud to call myself a Couch Grouser.
The gist is that this grouser, John Kosgrove, asked for an upgrade, citing the hassle and delay, and H. P. agreed.
It offers 16-inch and 20-inch track shoe options in single or triple grouser.
It is equipped with a maintenance-free crawler-type undercarriage with triple grouser plates or rubber crawlers.
Double grouser standard or narrow track shoes are available.
Due to customer demand in 2003 BLS was the first to develop and make a polyurethane bolt-on track pad that attaches to the steel triple grouser, the BLS Poly Bolt-On® TUFPADS® track pads.

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