What is a grouser?

What is a grouser? Here are some definitions.

Noun
  1. A person who grouses; a grumbler
  2. (nautical) A temporary pile driven into the bottom of a river to hold a boat in position; a spud
  3. (engineering) A protrusion on the surface of a wheel or continuous track segment, intended to increase traction in soil, snow, or other loose material
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Examples
It offers 16-inch and 20-inch track shoe options in single or triple grouser.
Founders of civilisation, interim riparian cotenants of the River Nile and purveyors of the Cheops Pyramid thrice, I am proud to call myself a Couch Grouser.
Malcolm turned, and found that the speaker was grouser Joliffe.
E wasn't arf a grouser, an' 'e 'ad good luck all the bloomin' time.

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