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What are doubletons?

What are doubletons? Here are some definitions.

Noun
  1. plural of doubleton
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A quatraton is inferred either from the presence of two singletons, no doubletons, and no tripletons or from the presence of no singletons, one doubleton, and no tripletons.
Five species occurred with more than 50 individuals, 15 species occurred with fewer than 10 individuals, two as doubletons, and two as singletons.
Of these, 75 are singletons, two are doubletons, and one is present in three sequences.
The distribution of singletons, doubletons, and other variants among the continents was investigated.

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