| We predict the relative fitnesses of hybrid genotypes by calculating the expected numbers of each type of incompatibility. |
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| We ignore environmental variation and deal directly with the fitnesses of genotypic values. |
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| But more importantly, it demonstrated that cultural evolution cannot be predicted purely on the basis of genetic fitnesses. |
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| We assume that the relative fitnesses of these genotypes remain constant. |
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| The differential fitnesses are weighted averages of phenotype fitnesses. |
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