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What is trituberculy?

What is trituberculy? Here are some definitions.

Noun
  1. (zoology) A theory of the development of mammalian molar teeth. The primitive stage is that of simple cones, as in reptiles. The simple cone then developed a smaller cone in front and another behind, and then a cingulum, and the three cones became arranged in a triangle (the trigon or trigonid; this stage is called the tritubercular or trigonodont), the two smaller cusps having moved to the outer side in upper and to the inner in lower molars. The trigon being a cutting apparatus, an extension of the posterior part of the crown was developed in lower molars for crushing, and a smaller corresponding part appeared in upper molars. Another large cone then arose, usually from the cingulum. In more complex forms, smaller intermediate cusps appeared.
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