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What are substages?

What are substages? Here are some definitions.

Noun
  1. plural of substage
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Examples
Fifth, to investigate the appropriateness of the ordering of TARPA substages, the current study conducted a statistical hierarchical analysis of the protocol.
Which, then, of these glacial substages provided the proper conditions and saw the first population of man in America?
The extraction combines six stages that could be broken down into some other substages.
Robe 11 and Robe III refer respectively to interstadial Substages Sa and Sc according to the terminology of Schwebel.
The respective frequencies of the leptotene, zygotene and pachytene stages as well as the frequencies of the four substages of pachytene were evaluated.
The 2002 AJCC pT2 substages confer no prognostic information on the rate of biochemical recurrence after radical prostatectomy.

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