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What are rank-and-filers?

What are rank-and-filers? Here are some definitions.

Noun
  1. plural of rank-and-filer
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Examples
Mr. Paterson, a former Senate minority leader, has limited tools to keep rank-and-filers in line.
Are all political rank-and-filers treated as cavalierly as we've been?
In opinion polls, Republican rank-and-filers, as distinct from Party professionals, strongly favor the idea of popular election.
The rank-and-filers think he does, and that's enough.
As Nixon hoped, white rank-and-filers abandoned the Democrats in droves.

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