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What is a progenitor?

What is a progenitor? Here are some definitions.

Noun
  1. A forefather, any of a person's direct ancestors.
  2. An individual from whom one or more people (dynasty, tribe, nation...) are descended.
  3. (biology) An ancestral form of a species
  4. (figuratively) A predecessor of something, especially if also a precursor or model.
  5. (figuratively) Someone who originates something.
  6. A founder.
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