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What are pinnules?

What are pinnules? Here are some definitions.

Noun
  1. plural of pinnule
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Examples
Predators mistaking the pinnule tips for the tegmen could take bites out of the distal parts of the pinnules.
The most common elements are parallel-veined leaves that resemble cordaites but that could be isolated pinnules of a pinnate leaf.
It has the same nodose and scaly root-stock, dark and polished stalk, glaucescent frond and mucronulate pinnules.
Fishelson and Nichols observed fish targeting organisms sheltering in the arms and pinnules of living crinoids.
Lepidopteris is a bipinnate frond whose pinnules resemble some species of Alethopteris.
The branchia is translucent white with brown spots and white-tipped brown pinnules.

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