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What are morphogens?

What are morphogens? Here are some definitions.

Noun
  1. plural of morphogen
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Examples
Hedgehog morphogens are associated with MPs shed from the plasma membrane of apoptotic stimulated T cells.
The Hedgehog proteins are secreted morphogens that provide positional information during the development of many multi-cellular animals.
The spread of their morphogens through epithelium promotes development.
The beautiful eyespots on butterfly wings are thought to rely on the principles described above, although involving more morphogens.
The researchers have thought that these so-called morphogens create chemical maps by simply diffusing through an embryo.
It's controlled by molecules called morphogens, which form concentration gradients along the head-to-tail axis, or other axes, of developing embryos.

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