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What are morphogens?

What are morphogens? Here are some definitions.

Noun
  1. plural of morphogen
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Examples
The Hedgehog proteins are secreted morphogens that provide positional information during the development of many multi-cellular animals.
He suggested that biological forms mirror patterns in the concentrations of hypothetical chemicals called morphogens.
The beautiful eyespots on butterfly wings are thought to rely on the principles described above, although involving more morphogens.
The spread of their morphogens through epithelium promotes development.
Hedgehog morphogens are associated with MPs shed from the plasma membrane of apoptotic stimulated T cells.
Plant growth and development is orchestrated by several classes of morphogens.

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