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What are keystones?

What are keystones? Here are some definitions.

Noun
  1. plural of keystone
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Examples
Risk management and evidence based medicine are keystones of clinical governance.
The keystones of the arches bear inscriptions indicating their precise location within the structure.
The building's exterior is largely unchanged, except that the carved mammal heads on the keystones of the first floor windows have been removed.
They can also question peers and learn how asking for and giving assistance to one another are keystones to academic success.
It is only after close study that apparent solecisms can be interpreted as the keystones of a highly conscious literary construct.
We must be careful, however, not to paint a picture of historical evidence that suggests that newspapers are the keystones of such documentation.

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