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What is an intergradation?

What is an intergradation? Here are some definitions.

Noun
  1. a gradual merging of two things
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Examples
Its eastern limit in Russia is hard to define, due to extensive hybridisation and intergradation with the Siberian spruce, but is usually given as the Ural Mountains.
Specimens from a transect of southeastern Sonora show intergradation between Perognathus goldmani and P. artus.
He called for deeper intergradation to achieve more unified GCC accomplishments.
No evidence of intergradation with any other race was noted.
Whether actual intergradation ever takes place across the narrow Bering Strait I do not know.
The extensive intergradation that has resulted from interbreeding is the chief cause of classification difficulties.

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