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What is a capitulum?

What is a capitulum? Here are some definitions.

Noun
  1. (botany) A densely clustered inflorescence composed of a large number of individual florets arising from a platform-like base.
  2. (arachnology) The head-like mouthpart apparatus of a tick, including the palpi, mandibles, and hypostome.
  3. (anatomy) A small protuberance on a bone which articulates into another bone to form a ball-and-socket joint.
  4. (entomology, obsolete) The enlarged end of a proboscis.
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As is well known, the ribs of the Monotremata have only the central head, the capitulum.
It is readily known by the numerous irregularly arranged scales at the base of the capitulum.
The achenes produced by each capitulum are similar, possess a pappus of bristles that causes them to move upwind and a well-developed elaisome.
The tuberculum and capitulum on each of the trunk ribs are separated only by a shallow concavity.
After fertilization, several achenes are produced per capitulum, and seeds are wind-dispersed.
It passes through the capitulum of the radius and the ulna and its coronoid process and radial notch.

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