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What is a barbule?

What is a barbule? Here are some definitions.

Noun
  1. (botany) A small barb or beard.
  2. (ornithology) Any of the secondary barbs that form a fringe of small projections on a feather.
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Examples
Even though there is a basic similarity with a flight feather, the peacock tail feather has an unusual barbule structure.
The scientists found that the outside layer of each barbule consisted of a two-dimensional crystal structure composed of melanin rods.
The animal residues found included possible blood residues, hair fragments, bone or antler fragments, and a feather barbule fragment.
Depending on how closely the rods were spaced and how many layers of rods were stacked up, the barbule could appear anything from yellow to green to blue to brown.
The jaw and lack of a chin barbule reveal the difference when look alike coalfish, a fellow member of the cod family, hunt for food in the same area.
Long spines are typically located on nodes of the mid-section of the barbule.

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