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What is a substratum?

What is a substratum? Here are some definitions.

Noun
  1. A layer that lies underneath another.
  2. (figuratively) The underlying cause or basis of something.
  3. (linguistics) A substrate.
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Examples
The downstream section was 50 to 60 m wide, straight, and substratum consisted of flocculent mud, 20 to 100 cm deep.
Rooted plants with three or four fully expanded leaves were transferred to a peat substratum and acclimatized in a glasshouse.
One was in contact with the substratum, and the other formed a dome within the marginal ossicles, thus creating a body cavity.
Such fishes can offset costs of transport using burst-and-glide behavior when swimming in the water column and ground effect near the substratum.
I am reluctant to state a case when the substratum was not the right substratum.
Serialization, for instance, has been attributed to substratum from the African languages, particularly the Kwa group.

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