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What are strongholds?

What are strongholds? Here are some definitions.

Noun
  1. plural of stronghold
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There were other strongholds at Dun Ara, and at Eilean Amalaig in Loch Spelve, where the MacLeans marshalled their birlinns.
It is also one of the last strongholds of the charismatic sage grouse, which has declined throughout the West.
The insurgents still had strongholds in the north-western district of Jolan, a warren of narrow streets.
These perches with huge vistas were just the sort of place the Urartians favoured for their strongholds.
Millions of pilgrims are expected to walk between Najaf and Karbala and other Shiite strongholds in Iraq.
Additional stocks of ammunition, food, medicines, and other materiel are created at strongholds.

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