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What are quills?

What are quills? Here are some definitions.

Noun
  1. plural of quill
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Examples
We were especially careful in excluding specimens in molt by checking all specimens for the presence of feather quills.
The clothing is made from white caribou hides and sewn with sinew, using split bird quills on the seams.
As is the case with North American porcupines, the quills are loosely attached but can't be thrown or otherwise projected.
Writers look to their quills, while painters care for horsehair and camel with as much care as palette and pigment.
And there it was, in plain sight by the door, with a small knife, the kind used to sharpen quills, hanging next to it.
The quills are so lightly fixed to the porcupine's body that they are easily detached and left imbedded in the attacker.

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