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What does growled mean?

Looking for the meaning or definition of the word growled? Here's what it means.

Verb
  1. simple past tense and past participle of growl
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Examples
The engine growled and the transmission grated, then settled down again as the truck started up a grade.
He then glared at Jake and growled, then charged him, his fist cocked back.
Jacks growled a bit, scowling, but Sam laughed, and smiled at him, so it melted away.
When he spoke his words were measured and careful, but they were growled at her.
I growled quietly under my breath and frowned at my reflection from a shop window.
I growled in frustration and barely restrained myself from ripping the next shirt I grabbed into pieces.

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