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What is a gemmule?

What is a gemmule? Here are some definitions.

Noun
  1. (biology) A small gemma or bud of dormant embryonic cells produced by some freshwater sponges
  2. (obsolete) A hypothetical particle once thought to be the basis of heredity
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Examples
The outer coat forms a kind of mantle by means of the skirts of which the gemmule is fastened to the support of the sponge.
The terminal growing bud of the axis is called the plumule or gemmule, and represents the ascending axis.
The effect of cadmium and mercury on gemmule formation and gemmosclere morphology in Ephydatia fluviatilis.
We can easily conceive a being so small, that a gemmule would be to it as large as St. Paul's would be to us.

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