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What does catenated mean?

Looking for the meaning or definition of the word catenated? Here's what it means.

Verb
  1. simple past tense and past participle of catenate
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Both the single-stranded circles and the double helical circles are catenated molecules.
As to segregation of replicated daughter molecules, to be discussed later, topoisomerase II is thought to locate DNA crossovers such as those present in catenated molecules.
These mutants lacked normal conidiophores and phialides and formed rod-shaped, catenated cells directly from hyphae.

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