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What is a testa?

What is a testa? Here are some definitions.

Noun
  1. (botany) A seed coat.
  2. (marine biology) The external calciferous shell, or endoskeleton, of an echinoderm; the test.
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Examples
Following the removal of the testa, the seed is cut along the line of the cotyledon suture.
Wheat grains consist of the seed coat, or testa, which surrounds the endosperm and embryo.
Sufficient moisture must be present to penetrate the testa or seed coat, but not so much that the seed rots or that the oxygen level in the soil is reduced.
While the subaleurone cells were intensively labelled, no label was seen in the outer layers of the grain, i.e. in the testa, pericarp and husk.
Seeds were scored daily for radicle emergence through the testa or PE envelope.
The seeds' hard testa is an effective protective structure allowing them to survive fire, and being passed through the guts of large animals which aid dispersal.

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