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What are substages?

What are substages? Here are some definitions.

Noun
  1. plural of substage
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Examples
The respective frequencies of the leptotene, zygotene and pachytene stages as well as the frequencies of the four substages of pachytene were evaluated.
The 2002 AJCC pT2 substages confer no prognostic information on the rate of biochemical recurrence after radical prostatectomy.
Fifth, to investigate the appropriateness of the ordering of TARPA substages, the current study conducted a statistical hierarchical analysis of the protocol.
Robe 11 and Robe III refer respectively to interstadial Substages Sa and Sc according to the terminology of Schwebel.
It represents the progress of disforestation since the early Neolithic and the presumed correspondence with the pattern of interglacial substages.
Plasma testosterone correlates with morph type across breeding substages in male White-throated Sparrows.

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