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What is a protoculture?

What is a protoculture? Here are some definitions.

Noun
  1. (physical anthropology) The passing of behaviours from one generation to another among non-human primates. These cultures are very rudimentary, and do not exhibit complex cultural technology.
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Protoculture supply was dwindling, the galaxywide war running down like clockwork as both sides' resources and infrastructures declined.

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