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What is a missing link?

What is a missing link? Here are some definitions.

Noun
  1. (colloquial) A hypothetical primate once thought necessary to explain a perceived evolutionary gap between apes and humans.
  2. (figuratively) Any sought-after or valuable intermediary figure or position.
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Examples
The missing link is the prosody, the patterns of stress, inflection, and intonation in a language.
Nonetheless, they provided the missing link that connected a flailing Wild West show tradition with the Western movie industry.
The purpose of this study is to provide one missing link in a growing chain of knowledge.
A dinosaur-dotty mum, who helped discover a missing link in our past, has been causing a real flutter in the scientific world.
In other words, these results provide the first direct evidence of the missing link between gas collision and the birth of the first stars.
Integration of these two data sets could provide a missing link in our understanding of the transition from aseismic to seismogenic plate motion.

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