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What are ligules?

What are ligules? Here are some definitions.

Noun
  1. plural of ligule
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Examples
They may represent tepals, staminodes, stipules, ligules, reduced bracts, leaves, branch systems, or novel structures.
Jagels and Garner observed the variation in callose deposition in the ligules of Selaginella.
Selection favoring recessives is common in maize breeding for several traits, such as sweetness, opacity, brachysm, lack of ligules.
The size of ligules were measured under light microscope and photographed with a Leica DMLB DC300 microphotographic microscope.

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