| The hard ticks are flattened dorsoventrally in the unfed state, possess a marginal outline which tapers toward the anterior, and the mouthparts are clearly visible. |
| The glenoid cavity is narrow and elongate dorsoventrally in the same plane as the limbs of the scapula. |
| While the femur could likely fold back against the side to a considerable degree, it cannot rotate dorsoventrally. |
| The distal end is dorsoventrally flattened with a mildly convex and faintly bilobed articulation surface. |
| The skull is dorsoventrally flattened, and partly disarticulated. |
| This clade is diagnosed by the dorsoventrally flattened fourth antennal segment and lack of sclerotization on the accessory scent gland. |
