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What are chuckles?

What are chuckles? Here are some definitions.

Noun
  1. plural of chuckle
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Examples
The Old Woman from the Royal Collection chuckles in her reverie, amused by the fantastic costume Rembrandt has garbed her in.
In spite of the physical demands of the production, however, the actors still found time to have a few chuckles on set.
You will be giving a lot of laughs, smiles, giggles, chuckles, hoots, snorts, cackle, titters, grins and guffaws.
Nick chuckles, and Jonnie guesses from the next muffled, unintelligible words that the pair are now kissing.
But it was hard to forget when every eddy of wind out of the east brought the smell of mud and the gleeful chuckles of the river.
He smiles and chuckles, plays happily with his toys and brings delight to everyone he meets.

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