| Antheridia formed on the dorsal and ventral surfaces of the wing, while archegonia formed on the ventral surface of the anterior cushion. |
| The necks of the archegonia hardly project above the general surface of the thallus. |
| The reproductive organs borne by the thallus or plant are called antheridia and archegonia, and serve for sexual reproduction. |
| Since the asexual gametophyte does not form antheridia or archegonia, crosses to confirm its genotype could not be performed. |
| The archegonia have a shorter neck than those of the ferns, and the neck is straight. |
| The archegonia are protected by being sunk in depressions of the disk or by a special two-lipped involucre. |
